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Functional Morphology - using biomechanics and manipulative experiments to understand how plant
reproductive structures work.
Cone Development - exploring the relationships between function, ontogeny, development, and the evolution of morphological diversity in conifers.
Cone Macroevolution - combining data from fossils, living species, and functional morphology to understand conifer cone evolution.
Phylogeny - using large molecular phylogenies to understand patterns of character evolution and their relationship to biogeography and diversification.
Paleobotany - finding and describing new fossil species to understand character evolution and morphological diversification in deep time .